maybe it's because you didn't erase on the lineart-layer but instead corrected lines with a white brush.
Just add another layer, paint bucket it a brightcolour and you'll probably see all the white-brush-spots from the lineart layer.
What you can do about it?
Either you try erase these white spots.
Redo the lineart.
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Or you could set the lineart layer to Multiply (select from the dropdown which probably has Normal right now in the Layer box) if you did it on Layer 1. ouo
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